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Studies in Drosophila have characterized Porcupine and Wntless to be essential for these steps, and their deficiency leads to phenotypes similar to those of Wingless (homolog of Wnt) mutants (van den Heuvel et al., 1993

(Source 3) 2

Long-term dosing studies have also been conducted to evaluate tissue tolerance and inflammation markers, suggesting a favorable profile in terms of local joint impact and systemic safety